Biological Sciences

Return to: USC Dornsife College of Letters, Arts and Sciences  

The Department of Biological Sciences is composed of four sections: human and evolutionary biology (HEB), marine and environmental biology (MEB), molecular and computational biology (MCB), and neurobiology (NBIO). A diversity of upper-division undergraduate and graduate courses enables students to gain a deep appreciation of organisms, from their molecular underpinnings to their interactions within ecosystems. We help students refine skills in critical thinking, communication, and collaboration, and develop an understanding of how biology can contribute to solving society’s challenges. We also offer students the opportunity to participate in the discovery of new knowledge by working alongside our faculty members in their research laboratories. Undergraduates in Biological Sciences have the opportunity to become involved in laboratory or field research and may enroll in the research courses BISC 290  or BISC 490x  for some of their elective units. Students who complete our degree programs will be well prepared for professional careers in the health sciences, as well as for careers in research and education in the basic biological and biomedical fields, and many other professions.

The department offers BA and BS degrees in Biological Sciences, and BA and BS degrees in Human Biology. See the Undergraduate Studies section of the HEB website for more information about the different tracks that are offered in the Human Biology undergraduate programs of study. The department also offers BA and BS degrees in Neuroscience (NEUR) and a BS in Computational Neuroscience (CPNS). See the Undergraduate Neuroscience program of the Dornsife website for descriptions of the undergraduate degrees in Neuroscience and Computational Neuroscience. For the BS degree in Biological Sciences, students who want a more focused curriculum may choose from four areas of emphasis: Biotechnology; Ecology, Evolution and Environment; Marine Biology; and Molecular, Cellular and Developmental Biology. The BS in Biochemistry is offered as a joint program with the Department of Chemistry. Minors are offered in Biology of Human Movement (HEB), Biology and Business (with the Marshall School of Business) (BISC), Craniofacial and Dental Technology (with the Herman Ostrow School of Dentistry) (BISC), Human Disease (HEB), Marine Biology (BISC), Natural Science (BISC) and Neuroscience (NEUR).

At the graduate level, the department offers challenging degree programs that lead to a PhD in Integrative and Evolutionary Biology (IEB), Marine Biology and Biological Oceanography (MEBP), and Molecular Biology (MB). See the Neurobiology section of the Dornsife website for a description of the graduate degree in Neuroscience (NGP). The department also offers MS and progressive MS degree programs in Marine and Environmental Biology and Molecular Genetics and Biochemistry.

Biological Sciences

Allan Hancock Foundation Building 107
(213) 740-1109
FAX: (213) 740-8123

Email (Undergraduate Programs): biodept@dornsife.usc.edu

Email (Graduate Programs): molecule@dornsife.usc.edu

Human and Evolutionary Biology

Location: Alan Hancock Foundation Building 254
(213) 821-9745

Email (Undergraduate Programs): d_heb@usc.edu

Email (Graduate Programs): molecule@dornsife.usc.edu

Department website: dornsife.usc.edu/bisc

Chair: Lorraine Turcotte, PhD

Section Heads

Human and Evolutionary Biology: Lorraine Turcotte, PhD

Marine and Environmental Biology: Douglas Capone, PhD

Molecular and Computational Biology: Oscar Aparicio, PhD

Neurobiology: Judith Hirsch, PhD

Faculty

President Emeritus, University Distinguished Chair in Science, Technology, and Society, Professor of Civil and Environmental Engineering, Professor of Population and Public Health Sciences and Professor of Biological Sciences: Carol L. Folt, PhD

USC Associates Captain Hancock Chair in Marine Science and Professor of Biological Sciences: David A. Caron, PhD*

Dana and David Dornsife Chair, Wrigley Institute Director and Professor of Psychology and Biological Sciences: Joseph Arvai, PhD (Psychology)

Harold Dornsife Chair in Neurosciences and Professor of Biological Sciences: Emily R. Liman, PhD

George and Louise Kawamoto Chair in Biological Sciences and Professor of Biological Sciences: David Hutchins, PhD

McCulloch-Crosby Chair in Marine Biology and Professor of Biological Sciences: Jed A. Fuhrman, PhD*

Rita and Edward Polusky Chair in Basic Cancer Research and Professor of Pathology and Biological Sciences: Michael R. Lieber, PhD (Pathology)

William and Julie Wrigley Chair in Environmental Studies and Professor of Biological Sciences: Douglas G. Capone, PhD

Wilford and Daris Zinsmeyer Early Career Chair in Marine Studies and Associate Professor of Biological Sciences: Carly Kenkel, PhD

University Professor and Milo Don and Lucille Appleman Professor of Biological Sciences and Professor of Biological Sciences and Psychology: Larry W. Swanson, PhD

Distinguished Professor of Biological Sciences: Susan L. Forsburg, PhD*

Provost Professor of Biological Sciences, Biomedical Engineering, Physiology and Biophysics, Stem Cell Biology and Regenerative Medicine, Pediatrics, Radiology, Ophthalmology and Quantitative and Computational Biology: Scott Fraser, PhD

Dean’s Professor of Biological Sciences and Professor of Biological Sciences, Medicine, Biomedical Engineering, Aerospace and Mechanical Engineering and Urology: Peter Kuhn, PhD

Gabilan Assistant Professor of Biological Sciences: Judith Kribelbauer, PhD

Gabilan Assistant Professor of Biological Sciences: Lauren McElvain, PhD

Gabilan Assistant Professor of Biological Sciences: Julia Schwartzman, PhD

Professors: Oscar M. Aparicio, PhD; Donald Arnold, PhD; David Bottjer, PhD (Earth Sciences); Sarah Bottjer, PhD; Lin Chen, PhD; Xiaojiang Chen, PhD; Pinchas Cohen, PhD (Gerontology); Dion Dickman, PhD; Casey Donovan, PhD; Suzanne Edmands, PhD; Ian Ehrenreich, PhD; Steven Finkel, PhD; Henryk Flashner, PhD (Aerospace and Mechanical Engineering); Carol L. Folt, PhD (Preventive Medicine); Myron F. Goodman, PhD; John F. Heidelberg, PhD; Albert A. Herrera, PhD*; Judith Hirsch, PhD; Scott Kanoski, PhD; Dale Kiefer, PhD; Naomi Levine, PhD; Donal T. Manahan, PhD; David D. McKemy, PhD*; Jill McNitt-Gray, PhD; Matthew Michael, PhD; James W. Moffett, PhD; Sergey Nuzhdin, PhD; Michael Quick, PhD*; David Raichlen, PhD; Sergio Sañudo-Wilhelmy, PhD; Craig Stanford, PhD; Cameron Thrash, PhD; John Tower, PhD; Lorraine Turcotte, PhD; Alan Watts, PhD; Eric Webb, PhD

Associate Professors: Irene Chiolo, PhD; Matthew Dean, PhD; Andrew Gracey, PhD; Bruce Herring, PhD; Samuel Andrew Hires, PhD; Rusty Lansford, PhD; Carolyn Phillips, PhD; Fabien Pinaud, PhD; Drew Steen, PhD

Assistant Professors: Tito Borner, PhD; Michael Campbell, PhD; Xianrui Cheng, PhD; Cornelius Gati, PhD; Noelle Held, PhD; Lauren McElvain, PhD; Jeffrey Moore, PhD; Derrick Morton, PhD; Lindsey Schier, PhD

Professors (Research): Feixue Fu, PhD; James Hicks, PhD

Professor (Teaching): Karla B. Heidelberg, PhD*

Professor (Clinical): Biren Patel, PhD (Integrative Anatomical Sciences)

Associate Professors (Research): Joel Hahn, PhD; Le Trinh, PhD

Associate Professors (Teaching): Christa Bancroft, PhD; Nancy Castro, PhD; Raffaella Ghittoni, PhD; Grayson Jaggers, PhD; Kurt Kwast, PhD; Gioia Polidori, PhD*; Trond Sigurdsen, PhD; Helen (Eleni) Soultanakis, PhD; Rory Spence, PhD

Associate Professor (Clinical): Kristi Lewton, PhD (Integrative Anatomical Sciences)

Assistant Professors (Research): Laura Gomez Consarnau, PhD; Phuong Pham, PhD; Thai Truong, PhD

Assistant Professors (Teaching): Cameron Egan, PhD; Chrisandra Kufeldt, PhD

Master Lecturer: Gudrun Floyd, MS

Lecturers: Rita Barakat, PhD; Joshua Carlos, MS; Charles (Kiki) Crawford, PhD; Shannon Cross, EdD; Laura Held, PhD; Lisa Le, MS; Bruce Teter, PhD

Adjunct Professors: Luis Chiappe, PhD; Gordon Hendler, PhD; Joel Martin, PhD; Witaya Mathiyakom, PhD; Tony Michaels, PhD

Adjunct Professors (Research): Kirk Fitzhugh, PhD; Myrna Jacobson, PhD; Xiaoming Wang, PhD

Adjunct Associate Professors (Research): Karen Kelly, PhD; Philip Requejo, PhD; Regina Wetzer, PhD

Adjunct Assistant Professors: Diane Kim, PhD; Benjamin Tully, PhD; JL Weissman, PhD

Adjunct Assistant Professors (Research): Andrew Jensen, PhD; Alberto Robador Ausejo, PhD; Emily Zakem, PhD

Adjunct Assistant Professors (Teaching): James Dines, PhD; Helaine Lopes, PhD

Emeritus: Michael Arbib, PhD; Norman Arnheim, PhD*; Richard Deonier, PhD; Linda Duguay, PhD; Caleb Finch, PhD; Dennis Hedgecock, PhD; Chien-Ping Ko, PhD; William O. McClure, PhD*; Andrew McMahon, PhD; Cornelius W Sullivan, PhD; Russel Zimmer, PhD

*Recipient of university-wide or college teaching award.

Undergraduate Degrees

Director of Undergraduate Studies, Biological Sciences: Cameron Egan, PhD

Director of Undergraduate Studies, Human Biology: Gioia Polidori, PhD

Director of Undergraduate Studies, Neuroscience: David McKemy, PhD

The undergraduate degree programs offered in Biological Sciences, Human Biology, and Neuroscience are designed for students passionate about biology and the natural sciences. Rigorous training in scientific methods paired with a broad liberal arts education prepares students completing these majors for careers in diverse fields such as business, education, law, health and technology. Many students choose the BISC, HBIO or Neuroscience programs because most of the course requirements overlap with the pre-health curriculum or prepare students for a career in academia or industry.

The department offers specially planned courses within the biological sciences to prepare students for admission to professional schools (medicine, dentistry, veterinary science, occupational therapy, physical therapy, pharmacy, optometry, public health), paramedical sciences (medical technology, physician’s assistant, clinical and public health microbiology, clinical biochemistry), naturalist and environmental positions in the public and private sectors, jobs in industry (biotechnology), and graduate study (basic biological and biomedical fields). With the proper selection of courses under the guidance of the Department of Biological Sciences and the USC Rossier School of Education, the BS degree satisfies the California requirements for secondary school teaching in the life sciences.

Students completing a degree in Human Biology [Human Biology (BA)  and Human Biology (BS) ] are expected to acquire the ability and skills to understand and interpret the human organism. Courses offer a variety of fields to examine humans, including anatomy, physiology, metabolism, neurobiology, genetics and behavior. Students will learn to conduct research in many different forms, communicate the outcome of an investigation, assess the contributions of research to the body of knowledge in Human Biology and understand how research data can be applied in an evolutionary context. To accomplish this outcome, our program provides state-of-the-art lab experiences with a focus on learner-centered, hands-on experiences using the latest web and computer technology, an extensive fossil collection of human ancestors along with many primate skeletons for the examination of human origins and their closest relatives in the Primate Order, opportunities for international experience via study-abroad programs and opportunities for high-level research in several areas, including the regulation of food intake, glucose intake and body weight, the mechanics and neural control of movement, and the behavioral ecology of humans, their ancestors, and nonhuman primates to understand evolutionary origins.

Advisement

Advisement in the Department of Biological Sciences is required each semester. First semester freshman and transfer advisement takes place during orientation. Advisement in all remaining semesters takes place during the pre-registration period. The USC Dornsife Office of Advising sends advisement appointment information each semester to all students enrolled in degree programs in Biological Sciences, Human Biology or Neuroscience.

Honors Program in Biological Sciences

The department offers an Honors Program to outstanding students already pursuing studies for the BA or BS degree in Biological Sciences or Human Biology. This program offers students an opportunity to participate in undergraduate research, experience in writing an honors thesis summarizing the completed research, and experience in an honors seminar. Honors students are required to take two semesters of  BISC 493x Honors Seminar  (1 unit/semester) and one semester of  BISC 494x Honors Thesis  (2 units) in addition to fulfilling all requirements of the BA or BS degree. Honors students must also choose  BISC 490x Directed Research  as one of their upper-division electives. This program leads to the designation on the transcript of Bachelor of Arts or Bachelor of Science in Biological Sciences or Human Biology with Honors.

Honors Admission Requirements

Students may apply to the department for admission to the Honors Program after having completed at least one year of work at USC with a minimum GPA of 3.5 in all science and math courses required for the major. Applications for the Honors Program are available in Allan Hancock Foundation (AHF), Room 105A or on the Biological Sciences website (dornsife.usc.edu/bisc).

Honors Scholarship Requirements

For continuation in the Honors Program, students must maintain a minimum GPA of 3.5 in the sciences and mathematics courses required for the major.

Honor Society

The Department of Biological Sciences offers membership in Phi Sigma, a national honor society, to selected biology majors (Alpha Alpha Chapter at USC.) Phi Sigma is devoted to the promotion of research and academic excellence in the biological sciences. Students with a GPA above 3.0 who have interest in research and have completed core requirements for the first two years in biological sciences are eligible. Major activities range from presentation of papers by members and lectures by outside speakers to field trips, laboratory demonstrations and joint research projects. 

Bachelor of Arts in Biological Sciences

This 56-unit degree program provides a solid foundation in the basic sciences: biology, biochemistry, molecular biology, genetics, chemistry, organic chemistry, calculus and physics. In addition, students complete 8 units of upper-division BISC elective courses. This degree program allows for more flexibility to pursue additional majors or minors, study abroad or graduate early.

Bachelor of Arts in Human Biology

The section of Human and Evolutionary Biology within the Department of Biological Sciences oversees the Human Biology BA at USC. This 45-unit degree is designed for students seeking a course of study in human applied physiology and metabolism, performance or human evolution. The Human Biology BA is an excellent option for students pursuing pre-professional education in the fields of physical therapy, nutrition, pharmacology, sports training, kinesiology, biomechanics, human evolution, primatology, evolutionary biology, etc.

Bachelor of Science in Biological Sciences

This 72-unit degree program includes the foundational science courses as the BA degree, but adds a statistics course requirement. Students in this degree program complete 20 units of upper-division elective courses. This degree program is excellent for students interested in research or graduate education in the natural sciences, and provides greater depth of study and opportunities to specialize.

Bachelor of Science in Human Biology

The section of Human and Evolutionary Biology within the Department of Biological Sciences oversees the Human Biology BS at USC. This 61-unit degree is designed for students seeking a focused course of study in human applied physiology, biomedicine, nutrition and metabolism or human evolution. The Human Biology BS is an excellent option for students pursuing pre-professional education in the fields of medicine, physical therapy, dentistry, physician assistant, nursing, nutrition, kinesiology, biomechanics, human evolution, primatology, evolutionary biology, etc.

Maymester Courses

Maymester courses are 4-week intensives that provide opportunities for students to engage in experiential learning off-campus. Maymester courses are held from mid-May to mid-June so that students can register for three courses during the traditional spring semester and one course in the Maymester. Maymester courses are part of the spring semester load, so tuition is charged as spring tuition.

Catalina Island

The Biological Sciences Department in conjunction with the USC Wrigley Institute for Environmental Studies (WIES) sponsors four-week Maymester programs at the USC Wrigley Marine Sciences Center (WMSC), on Santa Catalina Island. The Maymester courses focus on conservation biology, animal physiology, aquatic microbiology and field techniques in marine biology, and biological oceanography. These courses are open to all Biological Science majors as well as students in other departments and other institutions with a strong biology background. Students are primarily in their junior or senior years and may participate in any of the courses offered.

All the courses are taught by USC faculty and supported by USC graduate student teaching assistants. The classes are specialized to take advantage of the unique facilities and setting of Santa Catalina Island.

Students live on Catalina Island for the entire length of the course. Rates for room and board at the USC Wrigley Marine Science Center are comparable to those on campus. For those wishing to return to the mainland on the weekend — free transportation is provided each Friday to leave and each Monday to return.

For more information, students should contact the instructor(s) for the course of interest. Additional information can also be found at dornsife.usc.edu/flp/usc-wrigley-institute-on-catalina-island/.

Summer Faculty Led Programs - Problems Without Passports

Summer Faculty Led Program (FLP) courses are four-week intensives that provide opportunities for students to engage in experiential learning off-campus. Summer Faculty Led Program courses are held during the summer term. If you are interested in enrolling in an FLP course, please consult with Financial Aid to be sure that your award is available in time for the beginning of the summer session.  

Oxford, England

The Biological Sciences Department in conjunction with University of Oxford, Oxfordshire, England, sponsors a 4-week FLP course BISC 428 . This course studies illness and disease burdens that affects millions of people across the world. In addition, this course addresses: challenges in global health; health policy and public health; infectious diseases and chronic diseases; and international development and health.

Students will have the opportunity to learn in depth about health issues from practitioners in the field, who, throughout the course of their careers, have worked in various countries of Africa, South America, India and South East Asia. Students will also hear from researchers and clinicians active in vaccine development, disease burden abatement, chronic disease prevention, and international research trials. Together, the class will learn about the nuances and complexities that are woven into disease prevention and healthcare delivery around the world. A component of this course has students develop ideas for a sustainable solution to a current international health issue/problem.

For more information, students should contact the instructor of the course.

Minor in Biology and Business

The Marshall School of Business and the departments of Biological Sciences and Chemistry in the USC Dornsife College of Letters, Arts and Sciences jointly offer the minor in Biology and Business. See Biology and Business Minor  for a list of required courses.

Minor in the Biology of Human Movement

This minor provides students with an interdisciplinary perspective that integrates elements of anatomy, biomechanics, motor control as well as evolution and applies them to the assessment and improvement of human performance during the activities of daily living. This minor is ideal for students interested in fields such as bioengineering, biokinesiology, health promotion, athletics and dance.

Minor in Coaching, Health and Wellness

The minor in Coaching, Health and Wellness provides students with a foundation in exercise science, athletic coaching and principles of health and wellness.

Minor in Human Disease

The minor in Human Disease is an interdisciplinary program focused on the biological and psychological factors that determine wellness and disease. It is an ideal minor for students interested in various disciples such medicine, nursing, pharmacy, dentistry, public health, global health, health policy and health promotion. The minor includes two lower-division pre-requisites, two upper-division courses to be chosen from a list of HBIO courses and two upper-division courses to be chosen from a list of PSYC courses for a total of four upper-division courses (16 units).

Minor in Craniofacial and Dental Technology

For a description and complete listing of course requirements, see Craniofacial and Dental Technology Minor  in the Herman Ostrow School of Dentistry of USC section of the catalogue.

Minor in Marine Biology

This minor combines courses related specifically to marine science from several Dornsife College departments or programs. USC’s location along the Pacific Ocean and the unique facilities at the USC Wrigley Marine Science Center (WMSC) on Santa Catalina Island provide superb access for students to learn outside of traditional classroom venues. Students learn about the biology, evolution, and ecology of organisms that inhabit marine environments and the ecological and physical processes linking them. This program is an appropriate start for students who are seeking a focus in marine science or who are interested in broadening their base of knowledge about marine biology, earth sciences or environmental science.

Minor in Natural Science

The minor in natural science will first provide students with a foundation in the basic sciences of physics, chemistry and biology. Each student will then build on this foundation by selecting a variety of electives to meet individual scientific interests and academic goals. This minor is well-suited for pre-medical students who are non-science majors, or for students interested in developing a better understanding of the basic sciences.

This minor is not available to majors in the natural sciences or engineering.

Graduate Degrees

Progressive Master’s Degree Programs

Master of Science in Marine and Environmental Biology

The Master of Science degree in Marine and Environmental Biology (MEB) is designed to provide admitted students with a rigorous, quantitative and focused introduction to the burgeoning fields and breadth of topics in marine environmental biology/chemistry, geobiology, oceanography, conservation biology and population dynamics (depending upon the concentration selected). The MEB degree program provides students with independent research ex­periences that satisfy their own specific interests. The program is intended to position and stimulate students for possible advanced study leading to a PhD in one of the areas stated above, and/or provide a unique facet to the background of a prospective medical student. The program will also provide fundamental tools and expertise for entry into a master’s level position in academic, government, or private sector research laboratories. It will prepare students interested in governmental and non-government (NGO) environmental regulatory science and forge career pathways into private sector positions in environmental consulting and business. Details are available at dornsife.usc.edu/meb/.

Master of Science in Molecular Genetics and Biochemistry

This degree option is available for a limited number of highly qualified students who want to take an extra year for an intensive graduate-level research experience with Molecular Biology faculty. Students apply at the end of their junior year, generally after at least one semester of research in their proposed mentor’s laboratory. Application consists of a two-page research proposal, letter of support from the mentor, and transcript, which will be reviewed by the master’s committee for admission. Admission is not guaranteed. Students who are accepted will take the graduate core course in Molecular Biology (502a/b) as well as a literature seminar and relevant electives, but a substantial part of the credit will be graduate-level research. Students will be reviewed during their senior year to ensure progress. All MS students must complete a summative paper at the end of their master’s year to be approved by their mentor and one member of the master’s committee.

Doctoral Degree Programs in Biological Sciences

The graduate programs in biology provide education and training of biologists interested in living systems ranging from cellular to ecosystem levels of organization, investigated by laboratory or fieldwork. Courses and faculty research interests allow a multidisciplinary approach. A number of additional research areas are provided by adjunct faculty from other institutions, including the Los Angeles County Museum of Natural History and Children’s Hospital Los Angeles. Students develop the ability to formulate and test hypotheses, integrating information and concepts in the completion of a dissertation (PhD). A qualifying exam committee is formed for each student during the first year to develop a particular program of course work and research, and to evaluate the student’s progress. Specific information about the options in biological sciences can be obtained by requesting information brochures or online at dornsife.usc.edu/bisc.

Doctor of Philosophy in Integrative and Evolutionary Biology

This program of study provides each student with a broad, fundamental background in integrative and evolutionary biology (IEB) with in-depth specialization in one (or more) research areas. Broad research areas include, but are not limited to evolutionary biology, metabolism and nutrition, neuroscience, endocrinology, glycemic control, energy balance, biomechanics, biological anthropology, skeletal muscle physiology, kinesiology, and conservation. The curriculum and learning experiences for each student are tailored to their line of research. Students take a written and oral screening exam in the first year, and a written and oral qualifying exam no later than the end of the fifth semester of study.

This degree is awarded under the jurisdiction of the Graduate School. Refer to the Requirements for Graduation  section and The Graduate School  section of this catalogue for general regulations. All courses applied toward the degree must be courses accepted by the Graduate School.

Admission Requirements

Applicants must have a bachelor’s degree from an accredited four-year college or university, preferably in an area relevant to the research interests in IEB (e.g. biology, bioanthropology, biomedical engineering, psychology, etc.). Applicants are evaluated by their transcripts and GPA, three letters of recommendation, a statement of purpose and relevant research experience. A faculty member must serve as the sponsor of admission to the IEB program and adviser throughout the student’s course of study. We encourage applicants to reach out to potential faculty members in advance of submitting their application materials.

Financial Support

The program supports each student we admit in the form of teaching assistantships, research assistantships and fellowships. All awards include full tuition remission, a monthly stipend for living expenses, payment of student health/dental insurance and other university-wide benefits.

For any questions, please contact the IEB student services adviser.

Doctor of Philosophy in Marine Biology and Biological Oceanography

In the marine biology and biological oceanography (MBBO) program, each student receives a general background in marine sciences and obtains in-depth specialization in a research area of his or her choosing. Each student’s curriculum is fitted to the particular needs and demands of the chosen research field. Broad research areas include but are not limited to: organismal biology; evolution and population genetics; genomics; microbial biology, genetics and systems; aquatic, terrestrial and ocean ecology; environmental biology; and chemical biology. Typically, students take classes in the first two years. A written and oral qualifying exam to become a PhD candidate will be taken no later than the end of the fourth semester of study.

This degree is awarded under the jurisdiction of the Graduate School. Refer to the Requirements for Graduation  section and The Graduate School  section of this catalogue for general regulations. All courses applied toward the degrees must be courses accepted by the Graduate School.

Admission Requirements

Applicants must have a bachelor’s degree in a natural science (preferably biology) from an accredited four-year college or university. Undergraduate course work should include basic courses in biology, general physics, chemistry, organic chemistry, biochemistry and calculus. Applicants are evaluated by their transcripts and GPA, three letters of recommendation, and a statement of interest. A faculty member must serve as initial sponsor and adviser for admission to the MBBO program, so applicants are encouraged to contact potential advisers before applying. Applicants who are accepted into the program but judged to have minor deficiencies are expected to correct them within the first year.

Financial Support

The program supports each student we admit in the form of teaching assistantships, research assistantships and fellowships. All awards include full tuition remission, a monthly stipend for living expenses, payment of student health/dental insurance and other university-wide benefits.

For any questions, please contact the MBBO student services adviser.

Doctor of Philosophy in Molecular Biology

This program is designed to train the participants intensively in the concepts and experimental methodologies of molecular biology and biochemistry. The subject matter is organized in an integrated fashion (lectures, seminars and laboratory) to present fundamental information on the biochemistry, biophysics, genetics and development of cells from a variety of different organisms. Primary emphasis is on the relationship between structure and function at different integrative and functional levels. Molecular biology (MOL) students are required to complete at least three laboratory rotations in their first year.

This degree is awarded under the jurisdiction of the Graduate School. Refer to the Requirements for Graduation  section and The Graduate School  section of this catalogue for general regulations. All courses applied toward the degrees must be courses accepted by the Graduate School.

Admission Requirements

Application for admission may be accessed online at dornsife.usc.edu/mcb/phdhome/. Applicants are expected to have a bachelor’s degree or equivalent in a cognate area such as biology, chemistry, physics, engineering, bacteriology, computer science or bioinformatics. Undergraduate work should include a basic course in biology, basic physics, physical chemistry, organic chemistry, biochemistry and calculus from an accredited four-year college or university. Students who are deficient in any of these subjects may be required to correct the deficiency during the first two years of graduate study. Courses taken to correct these deficiencies may not be credited toward the degree. Applicants are evaluated holistically, through a combination of their transcripts, three letters of recommendation, and a statement of research interest. The letters of recommendation should be written by faculty members who can evaluate the promise of the student for graduate work and independent research.

Financial Support

The program supports each student we admit in the form of teaching assistantships, research assistantships and fellowships. All awards include full tuition remission, a monthly stipend for living expenses, payment of student health/dental insurance and other university-wide benefits.

For any questions, please contact the MOL student services adviser.

Programs

Bachelor’s Degree

• •  Biological Sciences (BA)
• •  Biological Sciences (Biotechnology) (BS)
• •  Biological Sciences (BS)
• •  Biological Sciences (Ecology, Evolution and Environment) (BS)
• •  Biological Sciences (Marine Biology) (BS)
• •  Biological Sciences (Molecular, Cellular and Developmental Biology) (BS)
• •  Human Biology (BA)
• •  Human Biology (BS)

Minor

• •  Biology and Business Minor
• •  Biology of Human Movement Minor
• •  Coaching, Health and Wellness Minor
• •  Human Disease Minor
• •  Marine Biology Minor
• •  Natural Science Minor

Master’s Degree

• •  Biology (MS)
• •  Marine and Environmental Biology (MS)
• •  Molecular and Computational Biology (MS)
• •  Molecular Genetics and Biochemistry (MS)

Doctoral Degree

• •  Biology (Marine and Environmental Biology) (PhD)
• •  Integrative and Evolutionary Biology (PhD)
• •  Marine Biology and Biological Oceanography (PhD)
• •  Molecular Biology (PhD)

Courses

Biological Sciences

• •  BISC 101Lgx Cellular and Molecular Biology
• •  BISC 102Lgx Humans and Their Environment
• •  BISC 103Lgx General Biology for the Environment and Life
• •  BISC 104Lgx How the Body Works
• •  BISC 108L Special Laboratory I
• •  BISC 109L Special Laboratory II
• •  BISC 110g Good Genes, Bad Genes
• •  BISC 112Lxg Data, Denial or Doom?: Talking about Climate Change
• •  BISC 115Lxg The Biology of Food
• •  BISC 120Lg General Biology: Organismal Biology and Evolution
• •  BISC 121Lg Advanced General Biology: Organismal Biology and Evolution
• •  BISC 140g Our Blue Planet in a Changing Climate
• •  BISC 150Lgx The Nature of Human Health and Disease
• •  BISC 180Lgx Evolution
• •  BISC 193 Introduction to Research I
• •  BISC 194 Introduction to Research II
• •  BISC 199 Neuroscience Colloquium
• •  BISC 220Lg General Biology: Cell Biology and Physiology
• •  BISC 221Lg Advanced General Biology: Cell Biology and Physiology
• •  BISC 230Lgx The Biology of the Brain
• •  BISC 235 The Biology of Sex
• •  BISC 290 Introduction to Biological Research
• •  BISC 300L Introduction to Microbiology
• •  BISC 303L The Kingdom Plantae: An Introduction to Botany
• •  BISC 305g Statistics for Biological Sciences
• •  BISC 307L General Physiology
• •  BISC 310 Statistical Thinking for Quantitative Biology
• •  BISC 312x Molecular Biochemistry
• •  BISC 313L Evolution and Population Genetics
• •  BISC 314L Cell Culture
• •  BISC 315L Introduction to Ecology
• •  BISC 320Lg Molecular Biology
• •  BISC 321 Multidisciplinary Seminar: Science, Technology and Society
• •  BISC 325 Genetics
• •  BISC 326L Urban Conservation Biology
• •  BISC 330L Biochemistry
• •  BISC 335 Science, Health and the Environment
• •  BISC 350 Bioethics
• •  BISC 352 Conservation Biology
• •  BISC 363L Mammalogy
• •  BISC 364L Advanced Techniques in Field Mammalogy
• •  BISC 365 Phylogenetics and Evolution
• •  BISC 369L Ecology and the Natural History of California
• •  BISC 371L Molecular Approaches to the Diversity of Life
• •  BISC 372 Cell Biology
• •  BISC 379L Our Future Changing Ocean
• •  BISC 390 Special Problems
• •  BISC 401 Introduction to Computational Analysis of Biological Data
• •  BISC 402 RNA Biology and Biotechnology
• •  BISC 403 Advanced Molecular Biology
• •  BISC 404L Ecology and Biodiversity
• •  BISC 405L General Embryology
• •  BISC 406L Biotechnology
• •  BISC 407 Cellular and Molecular Neuroscience
• •  BISC 408 Systems Neuroscience: From Synapses to Perception
• •  BISC 410 Applications of Molecular Biology to Medicine
• •  BISC 411 Advanced Cell Biology
• •  BISC 412 Oceans, Climate, and the Environment
• •  BISC 414 Biology of Cancer
• •  BISC 419L Microbiology for a Sustainable Future
• •  BISC 421 Neurobiology
• •  BISC 422L Neurobiology Laboratory
• •  BISC 423 Epilepsy to Ecstasy: Biological Basis of Neurological Disorders
• •  BISC 424 Brain Architecture
• •  BISC 425 Genetics through the Scientific Literature
• •  BISC 426 Principles of Neural Development
• •  BISC 427 The Global Environment
• •  BISC 428 The Biology of Health from a Global Perspective
• •  BISC 429 Cancer Immunology
• •  BISC 431L Aquatic Microbiology
• •  BISC 434 Introduction to Genome Science
• •  BISC 435 Advanced Biochemistry
• •  BISC 437L Comparative Physiology of Animals
• •  BISC 438 Nutritional Biochemistry
• •  BISC 440 Biodemography of Aging
• •  BISC 444 Practical Analysis of Biological Data in R
• •  BISC 445L Fundamentals of Vertebrate Biology
• •  BISC 447L Island Biogeography and Field Ecology
• •  BISC 448L Neuroanatomy
• •  BISC 450L Principles of Immunology
• •  BISC 451 Protein Engineering
• •  BISC 455L Molecular Approaches to Microbial Diversity — Catalina Semester
• •  BISC 456L Conservation Genetics
• •  BISC 457L Methods in Marine Biology and Biological Oceanography
• •  BISC 460 Seminar in Marine and Environmental Biology
• •  BISC 461 Seminar in Molecular and Computational Biology
• •  BISC 462 Seminar in Neurobiology
• •  BISC 466 Neuroimmunology
• •  BISC 469L Marine Biology
• •  BISC 473L Biological Oceanography
• •  BISC 474L Ecosystem Function and Earth Systems
• •  BISC 479L Computational Genome Analysis Laboratory
• •  BISC 480L Developmental Biology
• •  BISC 482 Systems Biology: Modeling the Dynamics of Life
• •  BISC 483 Geobiology and Astrobiology
• •  BISC 485 Advanced Seminar in Bacterial Survival and Evolution
• •  BISC 486 Regenerative Medicine: Principles, Paradigms and Practice
• •  BISC 487 Regenerative Medicine: Principles, Paradigms and Practice
• •  BISC 488 Microscopy and Spectroscopy of Biological Systems
• •  BISC 490x Directed Research